Pteridophyta are vascular plants that reproduce via spores rather than seeds. They include ferns, horsetails, and lycophytes. Pteridophytes were among the first plants to colonize land, having characteristics like a heterologous life cycle with a sporophyte generation bearing spores that develop into a gametophyte. They differ from other early land plants in having true roots, stems and leaves, as well as spores that form in sporangia on specialized sporophyll leaves.