Eurasian skylark

Appearance

The Eurasian skylark is 18–19 cm in length. Like most other larks, the Eurasian skylark is a rather dull-looking species, being mainly brown above and paler below. It has a short blunt crest on the head, which can be raised and lowered. In flight it shows a short tail and short broad wings. The tail and the rear edge of the wings are edged with white, which are visible when the bird is flying away, but not if it is heading towards the observer. The male has broader wings than the female. This adaptation for more efficient hovering flight may have evolved because of female Eurasian skylarks' preference for males that sing and hover for longer periods and so demonstrate that they are likely to have good overall fitness.

It is known for the song of the male, which is delivered in hovering flight from heights of 50 to 100 m, when the singing bird may appear as just a dot in the sky from the ground. The long, unbroken song is a clear, bubbling warble delivered high in the air while the bird is rising, circling or hovering. The song generally lasts two to three minutes, but it tends to last longer later in the mating season, when songs can last for 20 minutes or more. At wind farm sites, male skylarks have been found to sing at higher frequencies as a result of wind turbine noise.

Naming

Eleven subspecies are recognized:

⤷  "A. a. arvensis" Linnaeus, 1758 – northern, western and central Europe
⤷  "A. a. sierrae" Weigold, 1913 – Portugal, central and southern Spain
⤷  "A. a. harterti" Whitaker, 1904 – north-western Africa
⤷  "A. a. cantarella" Bonaparte, 1850 – southern Europe from north-eastern Spain to Turkey and the Caucasus
⤷  "A. a. armenica" Bogdanov, 1879 – south-eastern Turkey to Iran
⤷  "A. a. dulcivox" Hume, 1872 – south-eastern European Russia and western Siberia to north-western China and south-western Mongolia
⤷  "A. a. kiborti" Zaliesski, 1917 – southern Siberia, northern and eastern Mongolia and north-eastern China
⤷  "A. a. intermedia" R. Swinhoe, 1863 – north-central Siberia to north-eastern China and Korea
⤷  "A. a. pekinensis" Swinhoe, 1863 – north-eastern Siberia, Kamchatka Peninsula and the Kuril Islands
⤷  "A. a. lonnbergi" Hachisuka, 1926 – northern Sakhalin Island
⤷  "A. a. japonica" Temminck & Schlegel, 1848 – southern Sakhalin Island, southern Kuril Island, Japan and the Ryukyu Islands: the Japanese skylark

Some authorities recognise the subspecies "A. a. scotia" Tschusi, 1903 and "A. a. guillelmi" Witherby, 1921. In the above list "scotia" is included in the nominate subspecies "A. a. arvensis" and "guillelmi" is included in "A. a. sierrae".

Distribution

This lark breeds across most of Europe and Asia and in the mountains of north Africa. It is mainly resident in the west of its range, but eastern populations are more migratory, moving further south in winter. Even in the milder west of its range, many birds move to lowlands and the coast in winter. Asian birds, subspecies "A. a. pekinensis", appear as vagrants in Alaska.In the 19th century multiple batches of Eurasian skylarks were released in New Zealand beginning in 1864 in Nelson and in 1867 in Auckland. The wild population increased rapidly and had spread throughout both the North and South Islands by the 1920s.

In Australia the Eurasian skylark was introduced on multiple occasions beginning in 1850. It is now widespread in the southeast of the continent. In New South Wales it mostly occurs south of 33°S. It is widespread throughout Victoria and Tasmania and also occurs in the south-eastern corner of South Australia around Adelaide.

The Eurasian skylark was introduced to the southeastern Hawaiian Islands beginning in 1865. Although once common, it has declined in abundance on Oahu and is no longer found on Kauai. A study published in 1986 found European skylarks remained only on the islands of Hawaii and Maui and estimated a total population of 10,000 individuals.

The Eurasian skylark was introduced to Vancouver Island off the west coast of Canada in 1903; additional birds were introduced in 1913. The population grew and by 1962 there were around 1000 individuals. The numbers have subsequently declined due to loss of habitat, and in 2007 there were estimated to be only around 100 individuals spread over four small areas of the Saanich Peninsula.

Habitat

This lark breeds across most of Europe and Asia and in the mountains of north Africa. It is mainly resident in the west of its range, but eastern populations are more migratory, moving further south in winter. Even in the milder west of its range, many birds move to lowlands and the coast in winter. Asian birds, subspecies "A. a. pekinensis", appear as vagrants in Alaska.

Reproduction

Eurasian skylarks first breed when they are one year of age. Nesting may start in late March or early April. The nest is probably built by the female alone and is a shallow depression in the ground lined with grasses. The clutch is 3 to 5 eggs. The eggs of the nominate subspecies average 23.4 mm × 16.8 mm in size and weigh around 3.35 g. They have a grey-white or greenish background and are covered in brown or olive spots. They are incubated only by the female beginning after the last egg is laid and hatch synchronously after 11 days. The altricial young are cared for by both parents and for the first week are fed almost exclusively on insects. The nestlings fledge after 18 to 20 days but they usually leave the nest after 8 to 10 days. They are independent of their parents after around 25 days. The parents can have up to 4 broods in a season.

Food

The Eurasian skylark walks over the ground searching for food on the soil surface. Its diet consists of insects and plant material such as seeds and young leaves. Unlike a finch it swallows seeds without removing the husk. Insects form an important part of the diet in summer.

Predators

In the UK, Eurasian skylark numbers have declined over the last 30 years, as determined by the Common Bird Census started in the early 1960s by the British Trust for Ornithology. There are now only 10% of the numbers that were present 30 years ago. The RSPB have shown that this large decline is mainly due to changes in farming practices and only partly due to pesticides. In the past cereals were planted in the spring, grown through the summer and harvested in the early autumn. Cereals are now planted in the autumn, grown through the winter and are harvested in the early summer. The winter grown fields are much too dense in summer for the Eurasian skylark to be able to walk and run between the wheat stems to find its food.

A list of conservation interventions that could impact Alauda arvensis can be found in the Conservation Evidence website. English farmers are now encouraged and paid to maintain and create biodiversity for improving the habitat for Eurasian skylarks. Natural England's Environmental Stewardship Scheme offers 5 and 10-year grants for various beneficial options. For example, there is an option where the farmer can opt to grow a spring cereal instead of a winter one, and leave the stubble untreated with pesticide over the winter. The British Trust for Ornithology likens the stubbles to "giant bird tables" – providing spilt grain and weed seed to foraging birds.

The RSPB's research, over a six-year period, of winter-planted wheat fields has shown that suitable nesting areas for Eurasian skylarks can be made by turning the seeding machine off for a 5 to 10 metres stretch as the tractor goes over the ground to briefly stop the seeds being sown. This is repeated in several areas within the same field to make about two skylark plots per hectare. Subsequent spraying and fertilising can be continuous over the entire field. DEFRA suggests that Eurasian skylark plots should not be nearer than 24 m to the perimeter of the field, should not be near to telegraph poles, and should not be enclosed by trees.

When the crop grows, the Eurasian skylark plots become areas of low vegetation where Eurasian skylarks can easily hunt insects, and can build their well camouflaged ground nests. These areas of low vegetation are just right for skylarks, but the wheat in the rest of the field becomes too closely packed and too tall for the bird to seek food. At the RSPB's research farm in Cambridgeshire skylark numbers have increased threefold over six years. Fields where Eurasian skylarks were seen the year before would be obvious good sites for skylark plots. Farmers have reported that skylark plots are easy to make and the RSPB hope that this simple effective technique can be copied nationwide.

Cultural

When the word "lark" is used without specification, it usually refers to this species. A collective noun for Eurasian skylarks is an "exaltation". Although the "Oxford English Dictionary" describes this usage as "fanciful", it traces it back to a quotation from John Lydgate dating from about 1430. The verb "skylark", originally used by sailors, means "play tricks or practical jokes; indulge in horseplay, frolic". The verb and noun "lark", with similar meaning, may be related to "skylark" or to the dialect word "laik".

The bird is the subject of poems by Percy Bysshe Shelley, George Meredith, Ted Hughes, and numerous others; of a play by Henrik Ibsen entitled "A Doll's House" and of pieces of music including "The Lark Ascending" by Ralph Vaughan Williams. It is also the bird emblem of Kumamoto Prefecture. The Skylark of Space is a series of four science fiction novels by E.E. "Doc" Smith.